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Hedera helix

L. (1753)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Apiales Family: Araliaceae Genus: Hedera

English ivy

Description

Key Characters:

Growth Form: Woody vines, creeping or climbing by adventitious roots, secretory canals present in most parts.

Stems: Tips of young branches often pubescent with whitish 5–6–branched, stellate hairs.

Roots:

Leaves: Leaves simple. Alternate. Blades of sterile branches broadly ovate, 3–5–lobed, 4–10 cm long. those of fertile branches ovate to rhombic, entire. Base of leaves of sterile branches cordate. base of leaves of fertile branches obtuse to truncate. Margins shallowly lobed but otherwise entire. Petioles base often broad and sheathing the stem. Stipules absent.

Flowers: Flowers in simple terminal umbels or racemes of umbellules. Inflorescence often pubescent with whitish 5–6–branched, stellate hairs. Flowers bisexual (perfect). Pedicels often jointed below the ovary. Calyx a truncate rim or 5-toothed; often pubescent with whitish 5–6–branched, stellate hairs. Corolla of 5 petals, valvate. Stamens 5; alternate with the petals, distinct, inserted at the edge of the epigynous nectary disk. Ovary inferior, 5-carpellate, surmounted by a convex disk; carpels connate (rarely pseudomonomerous), with as many cells; ovules pendulous, 2 per cell, 1 abortive; styles as many as carpels, connate into a short stylopodium; stigmas sessile.

Fruit: Drupes dark purple at maturity; exocarp fleshy; pyrenes cartilaginous or membranous; often laterally compressed; globose; 5–8 mm long; the disk depressed–convex. Seeds 3–5 per fruit.

Ploidy: 2n = 43–48; 96

Habitat:

Elevation Range:

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Naturalized

Island Status

Kaua'i Naturalized
O'ahu Only found in cultivation
Maui Naturalized
Hawai'i Naturalized

Dispersal Agents

Birds

Vegetative Reproduction

Pollinators

Notes

  • Description digitized by Pumehana Imada
  • Native from Europe to the Caucasus Mountains, widely cultivated as an ornamental since ancient times; in Hawai‘i cultivated at least since the early 1900s and now very sparingly naturalized on Kaua‘i, O‘ahu, and Hawai‘i. Numerous horticultural forms and s

Bibliography

Name Published In: Sp. Pl.: 202 (1753)

Other References

Wagner et al. 1990:228 (K, O, H); Starr et al. 2003:24 (EM); Staples & Herbst 2005:133 (DESCR); Faccenda, Arthur, et al. 2025:156 (remove O [no evidence of nat.])

Wagner, W. L., Herbst, D. R., & Sohmer, S. H. (1999). Manual of the Flowering Plants of Hawai'i, Vols. 1 and 2 (No. Edn 2). University of Hawai'i and Bishop Museum Press.

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Hedera helix East Maui, Kula, old Kula Hwy PRESERVED_SPECIMEN Starr, F. Martz, K. Collector Number: 010419-1 Maui BISH 4/19/2001
2 Hedera helix captain cook; amy b. h. greenwell ethnobotanical gardens; abandoned coffee plantation at one side of garden In yard of house; sterile groundcover in deep shade & also climbing shrubs & tree trunks. PRESERVED_SPECIMEN herbst, d.r. staples iii, g.w. collector number: 9303 Hawaii BISH 1/11/1991
3 Hedera helix captain cook; amy b. h. greenwell ethnobotanical gardens Abandoned coffee plantation at one side of Garden; in yard of house. PRESERVED_SPECIMEN herbst, d.r. staples iii, g.w. collector number: 9303 Hawaii BISH 1/11/1991
4 Hedera helix east maui, kula, old kula hwy Along roadside with Syzygium cumini and Panicum maximum. PRESERVED_SPECIMEN starr, f. martz, k. collector number: 010419-1 Kauai BISH 4/19/2001
5 Hedera helix East Maui, Kula, old Kula Hwy PRESERVED_SPECIMEN Starr, F. 010419-1 Maui BISH 4/19/2001
6 Hedera helix W Maui, Lahaina District, Kaulalewelewe, near Haelaau Cabin PRESERVED_SPECIMEN H. Oppenheimer H40102 Maui HAPI 4/1/2001
7 Hedera helix 96746, Kapaa, HI, US Growing on and near the cabin. PRESERVED_SPECIMEN Oppenheimer, H. H40102 Maui HAPI 4/1/2001
8 Hedera helix East Maui, Kula, old Kula Hwy PRESERVED_SPECIMEN F. Starr & K. Martz 001218-1 Maui HAPI 12/18/2000
9 Hedera helix East Maui, Kula, old Kula Hwy Crawling on ground and climbing up trees. Associated vegetation Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN Starr, F.; Martz, K. 011026-1 Maui HAPI 10/26/2001
10 Hedera helix Oahu Island, Honolulu, University of Hawaii campus PRESERVED_SPECIMEN F. Starr & K. Martz 011026-1 Oahu HAPI 10/26/2001
11 Hedera helix East Maui, Makawao District, Piiholo, just below jct w. Olinda Rd Crawling over ground and climbing up trees to 10m ht. Associated vegetation: Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN Starr, F. 011026-1 Maui BISH 10/26/2001
12 Hedera helix honolulu; star market garden Purchased. PRESERVED_SPECIMEN staples, g.w. collector number: 951 Oahu BISH 7/19/1994
13 Hedera helix East Maui, Makawao District, at the top of Olinda Rd Climbing in Eucalyptus trees well beyond any obvious plantings. Associated vegetation: Pine plantations, Eucalyptus sp., Ulex europaeus, Kikuyu grass. PRESERVED_SPECIMEN Starr, F. Martz, K. Collector Number: 001218-1 Maui BISH 12/18/2000
14 Hedera helix honolulu; star market garden Purchased. PRESERVED_SPECIMEN staples, g.w. collector number: 952 Oahu BISH 7/19/1994
15 Hedera helix Waimea Canyon State Park; cultivated plants from cabins near Kokee Methodist Camp. Cultivated. PRESERVED_SPECIMEN Tim Flynn 2038 Kauai PTBG 12/30/1986
16 Hedera helix East Maui, Makawao District, Piiholo, just below jct w. Olinda Rd Crawling over ground and climbing up trees to 10m ht. Associated vegetation: Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN Starr, F. Martz, K. Collector Number: 011026-1 Maui BISH 10/26/2001
17 Hedera helix W Maui, Lahaina District, Kaulalewelewe, near Haelaau Cabin Growing on and near the cabin. PRESERVED_SPECIMEN Oppenheimer, H.L. H40102 Maui BISH 4/1/2001
18 Hedera helix east maui, makawao district, piiholo, just below jct w. olinda rd Crawling over ground and climbing up trees to 10m ht. Associated vegetation: Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN starr, f. martz, k. collector number: 011026-1 Maui BISH 10/26/2001
19 Hedera helix Hawaiian Islands, Maui, E maui, Piiholo, just below jct w. Olinda Rd. Crawling on ground and climbing up trees. Associated vegetation Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN Starr, F.; Martz, K. 011026-1 K 10/26/2001
20 Hedera helix puna; volcano In Degener's tree-fern forest. PRESERVED_SPECIMEN degener, o. degener, i. collector number: 32461 Hawaii BISH 7/23/1970
21 Hedera helix w maui, lahaina district, kaulalewelewe, near haelaau cabin PRESERVED_SPECIMEN oppenheimer, h.l. collector number: h40102 Kauai BISH 4/1/2001
22 Hedera helix Hawaiian Islands, Maui, W Maui, Lahaina District, Kaulalewelewe, near Haela'au Cabin. Growing on and near the cabin. PRESERVED_SPECIMEN Oppenheimer, H. H40102 K 4/1/2001
23 Hedera helix Kaulalewelewe. On and near Haela'au Cabin and spreading into Honokowai Valley and adjacent areas. PRESERVED_SPECIMEN Hank Oppenheimer H40102 Maui PTBG 4/1/2001
24 Hedera helix Kokee. Halemanu Road near Hedychium plantings. PRESERVED_SPECIMEN Tim Flynn 968 Kauai PTBG 12/11/1984
25 Hedera helix Hawaiian Islands, Maui, W Maui, Lahaina District, Kaulalewelewe, near Haela'au Cabin. PRESERVED_SPECIMEN H. Oppenheimer H40102 US 4/1/2001
26 Hedera helix E Maui, at the top of Olinda Rd PRESERVED_SPECIMEN F. Starr & K. Martz 001218-1 Maui US 12/18/2000
27 Hedera helix hawaiian islands, maui, w maui, lahaina district, kaulalewelewe, near haela'au cabin. PRESERVED_SPECIMEN h. oppenheimer h40102 US 4/1/2001
28 Hedera helix e maui, at the top of olinda rd PRESERVED_SPECIMEN f. starr & k. martz 001218-1 Maui US 12/18/2000
29 Hedera helix e maui, piiholo, just below jct w. olinda rd PRESERVED_SPECIMEN f. starr & k. martz 011026-1 Maui US 10/26/2001
30 Hedera helix E Maui, Piiholo, just below jct w. Olinda Rd PRESERVED_SPECIMEN Starr, F.; Martz, K. 011026-1 Maui US 10/26/2001
31 Hedera helix E Maui, Piiholo, just below jct w. Olinda Rd PRESERVED_SPECIMEN F. Starr & K. Martz 011026-1 Maui US 10/26/2001
32 Hedera helix E Maui, at the top of Olinda Rd PRESERVED_SPECIMEN Starr, F.; Martz, K. 001218-1 Maui US 12/18/2000
33 Hedera helix Hawaiian Islands, Maui, W Maui, Lahaina District, Kaulalewelewe, near Haela'au Cabin. PRESERVED_SPECIMEN Oppenheimer, H. H40102 US 4/1/2001
34 Hedera helix Puna; Volcano In Degener's tree-fern forest. PRESERVED_SPECIMEN Degener, O. 32461 Hawaii BISH 7/23/1970
35 Hedera helix Captain Cook; Amy B. H. Greenwell Ethnobotanical Gardens; abandoned coffee plantation at one side of Garden In yard of house; in deep shade & also climbing shrubs & tree trunks. PRESERVED_SPECIMEN Herbst, D.R. 9303 Hawaii BISH 1/11/1991
36 Hedera helix East Maui, Makawao District, at the top of Olinda Rd Climbing in Eucalyptus trees well beyond any obvious plantings. Associated vegetation: Pine plantations, Eucalyptus sp., Ulex europaeus, Kikuyu grass. PRESERVED_SPECIMEN Starr, F. 001218-1 Maui BISH 12/18/2000
37 Hedera helix W Maui, Lahaina District, Kaulalewelewe, near Haelaau Cabin Growing on and near the cabin. PRESERVED_SPECIMEN Oppenheimer, H.L. Collector Number: H40102 Maui BISH 4/1/2001
38 Hedera helix Honolulu; Star Market Garden Purchased. PRESERVED_SPECIMEN Staples, G. 952 Oahu BISH 7/19/1994
39 Hedera helix Honolulu; Star Market Garden Purchased. PRESERVED_SPECIMEN Staples, G. 951 Oahu BISH 7/19/1994
40 Hedera helix east maui, makawao district, at the top of olinda rd Climbing in Eucalyptus trees well beyond any obvious plantings. Associated vegetation: Pine plantations, Eucalyptus sp., Ulex europaeus, Kikuyu grass. PRESERVED_SPECIMEN starr, f. martz, k. collector number: 001218-1 Maui BISH 12/18/2000
41 Hedera helix east maui, kula, old kula hwy PRESERVED_SPECIMEN starr, f. martz, k. collector number: 010419-1 Oahu BISH 4/19/2001
42 Hedera helix East Maui, Kula, old Kula Hwy PRESERVED_SPECIMEN Starr, F. Martz, K. Collector Number: 010419-1 Maui BISH 4/19/2001
43 Hedera helix captain cook; amy b. h. greenwell ethnobotanical gardens; abandoned coffee plantation at one side of garden In yard of house; sterile groundcover in deep shade & also climbing shrubs & tree trunks. PRESERVED_SPECIMEN herbst, d.r. staples iii, g.w. collector number: 9303 Hawaii BISH 1/11/1991
44 Hedera helix captain cook; amy b. h. greenwell ethnobotanical gardens Abandoned coffee plantation at one side of Garden; in yard of house. PRESERVED_SPECIMEN herbst, d.r. staples iii, g.w. collector number: 9303 Hawaii BISH 1/11/1991
45 Hedera helix east maui, kula, old kula hwy Along roadside with Syzygium cumini and Panicum maximum. PRESERVED_SPECIMEN starr, f. martz, k. collector number: 010419-1 Kauai BISH 4/19/2001
46 Hedera helix East Maui, Kula, old Kula Hwy PRESERVED_SPECIMEN Starr, F. 010419-1 Maui BISH 4/19/2001
47 Hedera helix W Maui, Lahaina District, Kaulalewelewe, near Haelaau Cabin PRESERVED_SPECIMEN H. Oppenheimer H40102 Maui HAPI 4/1/2001
48 Hedera helix 96746, Kapaa, HI, US Growing on and near the cabin. PRESERVED_SPECIMEN Oppenheimer, H. H40102 Maui HAPI 4/1/2001
49 Hedera helix East Maui, Kula, old Kula Hwy PRESERVED_SPECIMEN F. Starr & K. Martz 001218-1 Maui HAPI 12/18/2000
50 Hedera helix East Maui, Kula, old Kula Hwy Crawling on ground and climbing up trees. Associated vegetation Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN Starr, F.; Martz, K. 011026-1 Maui HAPI 10/26/2001
51 Hedera helix Oahu Island, Honolulu, University of Hawaii campus PRESERVED_SPECIMEN F. Starr & K. Martz 011026-1 Oahu HAPI 10/26/2001
52 Hedera helix East Maui, Makawao District, Piiholo, just below jct w. Olinda Rd Crawling over ground and climbing up trees to 10m ht. Associated vegetation: Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN Starr, F. 011026-1 Maui BISH 10/26/2001
53 Hedera helix honolulu; star market garden Purchased. PRESERVED_SPECIMEN staples, g.w. collector number: 951 Oahu BISH 7/19/1994
54 Hedera helix East Maui, Makawao District, at the top of Olinda Rd Climbing in Eucalyptus trees well beyond any obvious plantings. Associated vegetation: Pine plantations, Eucalyptus sp., Ulex europaeus, Kikuyu grass. PRESERVED_SPECIMEN Starr, F. Martz, K. Collector Number: 001218-1 Maui BISH 12/18/2000
55 Hedera helix honolulu; star market garden Purchased. PRESERVED_SPECIMEN staples, g.w. collector number: 952 Oahu BISH 7/19/1994
56 Hedera helix Waimea Canyon State Park; cultivated plants from cabins near Kokee Methodist Camp. Cultivated. PRESERVED_SPECIMEN Tim Flynn 2038 Kauai PTBG 12/30/1986
57 Hedera helix East Maui, Makawao District, Piiholo, just below jct w. Olinda Rd Crawling over ground and climbing up trees to 10m ht. Associated vegetation: Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN Starr, F. Martz, K. Collector Number: 011026-1 Maui BISH 10/26/2001
58 Hedera helix W Maui, Lahaina District, Kaulalewelewe, near Haelaau Cabin Growing on and near the cabin. PRESERVED_SPECIMEN Oppenheimer, H.L. H40102 Maui BISH 4/1/2001
59 Hedera helix east maui, makawao district, piiholo, just below jct w. olinda rd Crawling over ground and climbing up trees to 10m ht. Associated vegetation: Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN starr, f. martz, k. collector number: 011026-1 Maui BISH 10/26/2001
60 Hedera helix Hawaiian Islands, Maui, E maui, Piiholo, just below jct w. Olinda Rd. Crawling on ground and climbing up trees. Associated vegetation Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN Starr, F.; Martz, K. 011026-1 K 10/26/2001
61 Hedera helix puna; volcano In Degener's tree-fern forest. PRESERVED_SPECIMEN degener, o. degener, i. collector number: 32461 Hawaii BISH 7/23/1970
62 Hedera helix w maui, lahaina district, kaulalewelewe, near haelaau cabin PRESERVED_SPECIMEN oppenheimer, h.l. collector number: h40102 Kauai BISH 4/1/2001
63 Hedera helix Hawaiian Islands, Maui, W Maui, Lahaina District, Kaulalewelewe, near Haela'au Cabin. Growing on and near the cabin. PRESERVED_SPECIMEN Oppenheimer, H. H40102 K 4/1/2001
64 Hedera helix Kaulalewelewe. On and near Haela'au Cabin and spreading into Honokowai Valley and adjacent areas. PRESERVED_SPECIMEN Hank Oppenheimer H40102 Maui PTBG 4/1/2001
65 Hedera helix Kokee. Halemanu Road near Hedychium plantings. PRESERVED_SPECIMEN Tim Flynn 968 Kauai PTBG 12/11/1984
66 Hedera helix Hawaiian Islands, Maui, W Maui, Lahaina District, Kaulalewelewe, near Haela'au Cabin. PRESERVED_SPECIMEN H. Oppenheimer H40102 US 4/1/2001
67 Hedera helix E Maui, at the top of Olinda Rd PRESERVED_SPECIMEN F. Starr & K. Martz 001218-1 Maui US 12/18/2000
68 Hedera helix hawaiian islands, maui, w maui, lahaina district, kaulalewelewe, near haela'au cabin. PRESERVED_SPECIMEN h. oppenheimer h40102 US 4/1/2001
69 Hedera helix e maui, at the top of olinda rd PRESERVED_SPECIMEN f. starr & k. martz 001218-1 Maui US 12/18/2000
70 Hedera helix e maui, piiholo, just below jct w. olinda rd PRESERVED_SPECIMEN f. starr & k. martz 011026-1 Maui US 10/26/2001
71 Hedera helix E Maui, Piiholo, just below jct w. Olinda Rd PRESERVED_SPECIMEN Starr, F.; Martz, K. 011026-1 Maui US 10/26/2001
72 Hedera helix E Maui, Piiholo, just below jct w. Olinda Rd PRESERVED_SPECIMEN F. Starr & K. Martz 011026-1 Maui US 10/26/2001
73 Hedera helix E Maui, at the top of Olinda Rd PRESERVED_SPECIMEN Starr, F.; Martz, K. 001218-1 Maui US 12/18/2000
74 Hedera helix Hawaiian Islands, Maui, W Maui, Lahaina District, Kaulalewelewe, near Haela'au Cabin. PRESERVED_SPECIMEN Oppenheimer, H. H40102 US 4/1/2001
75 Hedera helix Puna; Volcano In Degener's tree-fern forest. PRESERVED_SPECIMEN Degener, O. 32461 Hawaii BISH 7/23/1970
76 Hedera helix Captain Cook; Amy B. H. Greenwell Ethnobotanical Gardens; abandoned coffee plantation at one side of Garden In yard of house; in deep shade & also climbing shrubs & tree trunks. PRESERVED_SPECIMEN Herbst, D.R. 9303 Hawaii BISH 1/11/1991
77 Hedera helix East Maui, Makawao District, at the top of Olinda Rd Climbing in Eucalyptus trees well beyond any obvious plantings. Associated vegetation: Pine plantations, Eucalyptus sp., Ulex europaeus, Kikuyu grass. PRESERVED_SPECIMEN Starr, F. 001218-1 Maui BISH 12/18/2000
78 Hedera helix W Maui, Lahaina District, Kaulalewelewe, near Haelaau Cabin Growing on and near the cabin. PRESERVED_SPECIMEN Oppenheimer, H.L. Collector Number: H40102 Maui BISH 4/1/2001
79 Hedera helix Honolulu; Star Market Garden Purchased. PRESERVED_SPECIMEN Staples, G. 952 Oahu BISH 7/19/1994
80 Hedera helix Honolulu; Star Market Garden Purchased. PRESERVED_SPECIMEN Staples, G. 951 Oahu BISH 7/19/1994
81 Hedera helix east maui, makawao district, at the top of olinda rd Climbing in Eucalyptus trees well beyond any obvious plantings. Associated vegetation: Pine plantations, Eucalyptus sp., Ulex europaeus, Kikuyu grass. PRESERVED_SPECIMEN starr, f. martz, k. collector number: 001218-1 Maui BISH 12/18/2000
82 Hedera helix east maui, kula, old kula hwy PRESERVED_SPECIMEN starr, f. martz, k. collector number: 010419-1 Oahu BISH 4/19/2001
83 Hedera helix East Maui, Kula, old Kula Hwy PRESERVED_SPECIMEN Starr, F. Martz, K. Collector Number: 010419-1 Maui BISH 4/19/2001
84 Hedera helix captain cook; amy b. h. greenwell ethnobotanical gardens; abandoned coffee plantation at one side of garden In yard of house; sterile groundcover in deep shade & also climbing shrubs & tree trunks. PRESERVED_SPECIMEN herbst, d.r. staples iii, g.w. collector number: 9303 Hawaii BISH 1/11/1991
85 Hedera helix captain cook; amy b. h. greenwell ethnobotanical gardens Abandoned coffee plantation at one side of Garden; in yard of house. PRESERVED_SPECIMEN herbst, d.r. staples iii, g.w. collector number: 9303 Hawaii BISH 1/11/1991
86 Hedera helix east maui, kula, old kula hwy Along roadside with Syzygium cumini and Panicum maximum. PRESERVED_SPECIMEN starr, f. martz, k. collector number: 010419-1 Kauai BISH 4/19/2001
87 Hedera helix East Maui, Kula, old Kula Hwy PRESERVED_SPECIMEN Starr, F. 010419-1 Maui BISH 4/19/2001
88 Hedera helix W Maui, Lahaina District, Kaulalewelewe, near Haelaau Cabin PRESERVED_SPECIMEN H. Oppenheimer H40102 Maui HAPI 4/1/2001
89 Hedera helix 96746, Kapaa, HI, US Growing on and near the cabin. PRESERVED_SPECIMEN Oppenheimer, H. H40102 Maui HAPI 4/1/2001
90 Hedera helix East Maui, Kula, old Kula Hwy PRESERVED_SPECIMEN F. Starr & K. Martz 001218-1 Maui HAPI 12/18/2000
91 Hedera helix East Maui, Kula, old Kula Hwy Crawling on ground and climbing up trees. Associated vegetation Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN Starr, F.; Martz, K. 011026-1 Maui HAPI 10/26/2001
92 Hedera helix Oahu Island, Honolulu, University of Hawaii campus PRESERVED_SPECIMEN F. Starr & K. Martz 011026-1 Oahu HAPI 10/26/2001
93 Hedera helix East Maui, Makawao District, Piiholo, just below jct w. Olinda Rd Crawling over ground and climbing up trees to 10m ht. Associated vegetation: Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN Starr, F. 011026-1 Maui BISH 10/26/2001
94 Hedera helix honolulu; star market garden Purchased. PRESERVED_SPECIMEN staples, g.w. collector number: 951 Oahu BISH 7/19/1994
95 Hedera helix East Maui, Makawao District, at the top of Olinda Rd Climbing in Eucalyptus trees well beyond any obvious plantings. Associated vegetation: Pine plantations, Eucalyptus sp., Ulex europaeus, Kikuyu grass. PRESERVED_SPECIMEN Starr, F. Martz, K. Collector Number: 001218-1 Maui BISH 12/18/2000
96 Hedera helix honolulu; star market garden Purchased. PRESERVED_SPECIMEN staples, g.w. collector number: 952 Oahu BISH 7/19/1994
97 Hedera helix Waimea Canyon State Park; cultivated plants from cabins near Kokee Methodist Camp. Cultivated. PRESERVED_SPECIMEN Tim Flynn 2038 Kauai PTBG 12/30/1986
98 Hedera helix East Maui, Makawao District, Piiholo, just below jct w. Olinda Rd Crawling over ground and climbing up trees to 10m ht. Associated vegetation: Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN Starr, F. Martz, K. Collector Number: 011026-1 Maui BISH 10/26/2001
99 Hedera helix W Maui, Lahaina District, Kaulalewelewe, near Haelaau Cabin Growing on and near the cabin. PRESERVED_SPECIMEN Oppenheimer, H.L. H40102 Maui BISH 4/1/2001
100 Hedera helix east maui, makawao district, piiholo, just below jct w. olinda rd Crawling over ground and climbing up trees to 10m ht. Associated vegetation: Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN starr, f. martz, k. collector number: 011026-1 Maui BISH 10/26/2001
101 Hedera helix Hawaiian Islands, Maui, E maui, Piiholo, just below jct w. Olinda Rd. Crawling on ground and climbing up trees. Associated vegetation Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN Starr, F.; Martz, K. 011026-1 K 10/26/2001
102 Hedera helix puna; volcano In Degener's tree-fern forest. PRESERVED_SPECIMEN degener, o. degener, i. collector number: 32461 Hawaii BISH 7/23/1970
103 Hedera helix w maui, lahaina district, kaulalewelewe, near haelaau cabin PRESERVED_SPECIMEN oppenheimer, h.l. collector number: h40102 Kauai BISH 4/1/2001
104 Hedera helix Hawaiian Islands, Maui, W Maui, Lahaina District, Kaulalewelewe, near Haela'au Cabin. Growing on and near the cabin. PRESERVED_SPECIMEN Oppenheimer, H. H40102 K 4/1/2001
105 Hedera helix Kaulalewelewe. On and near Haela'au Cabin and spreading into Honokowai Valley and adjacent areas. PRESERVED_SPECIMEN Hank Oppenheimer H40102 Maui PTBG 4/1/2001
106 Hedera helix Kokee. Halemanu Road near Hedychium plantings. PRESERVED_SPECIMEN Tim Flynn 968 Kauai PTBG 12/11/1984
107 Hedera helix Hawaiian Islands, Maui, W Maui, Lahaina District, Kaulalewelewe, near Haela'au Cabin. PRESERVED_SPECIMEN H. Oppenheimer H40102 US 4/1/2001
108 Hedera helix E Maui, at the top of Olinda Rd PRESERVED_SPECIMEN F. Starr & K. Martz 001218-1 Maui US 12/18/2000
109 Hedera helix hawaiian islands, maui, w maui, lahaina district, kaulalewelewe, near haela'au cabin. PRESERVED_SPECIMEN h. oppenheimer h40102 US 4/1/2001
110 Hedera helix e maui, at the top of olinda rd PRESERVED_SPECIMEN f. starr & k. martz 001218-1 Maui US 12/18/2000
111 Hedera helix e maui, piiholo, just below jct w. olinda rd PRESERVED_SPECIMEN f. starr & k. martz 011026-1 Maui US 10/26/2001
112 Hedera helix E Maui, Piiholo, just below jct w. Olinda Rd PRESERVED_SPECIMEN Starr, F.; Martz, K. 011026-1 Maui US 10/26/2001
113 Hedera helix E Maui, Piiholo, just below jct w. Olinda Rd PRESERVED_SPECIMEN F. Starr & K. Martz 011026-1 Maui US 10/26/2001
114 Hedera helix E Maui, at the top of Olinda Rd PRESERVED_SPECIMEN Starr, F.; Martz, K. 001218-1 Maui US 12/18/2000
115 Hedera helix Hawaiian Islands, Maui, W Maui, Lahaina District, Kaulalewelewe, near Haela'au Cabin. PRESERVED_SPECIMEN Oppenheimer, H. H40102 US 4/1/2001
116 Hedera helix Puna; Volcano In Degener's tree-fern forest. PRESERVED_SPECIMEN Degener, O. 32461 Hawaii BISH 7/23/1970
117 Hedera helix Captain Cook; Amy B. H. Greenwell Ethnobotanical Gardens; abandoned coffee plantation at one side of Garden In yard of house; in deep shade & also climbing shrubs & tree trunks. PRESERVED_SPECIMEN Herbst, D.R. 9303 Hawaii BISH 1/11/1991
118 Hedera helix East Maui, Makawao District, at the top of Olinda Rd Climbing in Eucalyptus trees well beyond any obvious plantings. Associated vegetation: Pine plantations, Eucalyptus sp., Ulex europaeus, Kikuyu grass. PRESERVED_SPECIMEN Starr, F. 001218-1 Maui BISH 12/18/2000
119 Hedera helix W Maui, Lahaina District, Kaulalewelewe, near Haelaau Cabin Growing on and near the cabin. PRESERVED_SPECIMEN Oppenheimer, H.L. Collector Number: H40102 Maui BISH 4/1/2001
120 Hedera helix Honolulu; Star Market Garden Purchased. PRESERVED_SPECIMEN Staples, G. 952 Oahu BISH 7/19/1994
121 Hedera helix Honolulu; Star Market Garden Purchased. PRESERVED_SPECIMEN Staples, G. 951 Oahu BISH 7/19/1994
122 Hedera helix east maui, makawao district, at the top of olinda rd Climbing in Eucalyptus trees well beyond any obvious plantings. Associated vegetation: Pine plantations, Eucalyptus sp., Ulex europaeus, Kikuyu grass. PRESERVED_SPECIMEN starr, f. martz, k. collector number: 001218-1 Maui BISH 12/18/2000
123 Hedera helix east maui, kula, old kula hwy PRESERVED_SPECIMEN starr, f. martz, k. collector number: 010419-1 Oahu BISH 4/19/2001
124 Hedera helix East Maui, Kula, old Kula Hwy PRESERVED_SPECIMEN Starr, F. Martz, K. Collector Number: 010419-1 Maui BISH 4/19/2001
125 Hedera helix captain cook; amy b. h. greenwell ethnobotanical gardens; abandoned coffee plantation at one side of garden In yard of house; sterile groundcover in deep shade & also climbing shrubs & tree trunks. PRESERVED_SPECIMEN herbst, d.r. staples iii, g.w. collector number: 9303 Hawaii BISH 1/11/1991
126 Hedera helix captain cook; amy b. h. greenwell ethnobotanical gardens Abandoned coffee plantation at one side of Garden; in yard of house. PRESERVED_SPECIMEN herbst, d.r. staples iii, g.w. collector number: 9303 Hawaii BISH 1/11/1991
127 Hedera helix east maui, kula, old kula hwy Along roadside with Syzygium cumini and Panicum maximum. PRESERVED_SPECIMEN starr, f. martz, k. collector number: 010419-1 Kauai BISH 4/19/2001
128 Hedera helix East Maui, Kula, old Kula Hwy PRESERVED_SPECIMEN Starr, F. 010419-1 Maui BISH 4/19/2001
129 Hedera helix W Maui, Lahaina District, Kaulalewelewe, near Haelaau Cabin PRESERVED_SPECIMEN H. Oppenheimer H40102 Maui HAPI 4/1/2001
130 Hedera helix 96746, Kapaa, HI, US Growing on and near the cabin. PRESERVED_SPECIMEN Oppenheimer, H. H40102 Maui HAPI 4/1/2001
131 Hedera helix East Maui, Kula, old Kula Hwy PRESERVED_SPECIMEN F. Starr & K. Martz 001218-1 Maui HAPI 12/18/2000
132 Hedera helix East Maui, Kula, old Kula Hwy Crawling on ground and climbing up trees. Associated vegetation Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN Starr, F.; Martz, K. 011026-1 Maui HAPI 10/26/2001
133 Hedera helix Oahu Island, Honolulu, University of Hawaii campus PRESERVED_SPECIMEN F. Starr & K. Martz 011026-1 Oahu HAPI 10/26/2001
134 Hedera helix East Maui, Makawao District, Piiholo, just below jct w. Olinda Rd Crawling over ground and climbing up trees to 10m ht. Associated vegetation: Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN Starr, F. 011026-1 Maui BISH 10/26/2001
135 Hedera helix honolulu; star market garden Purchased. PRESERVED_SPECIMEN staples, g.w. collector number: 951 Oahu BISH 7/19/1994
136 Hedera helix East Maui, Makawao District, at the top of Olinda Rd Climbing in Eucalyptus trees well beyond any obvious plantings. Associated vegetation: Pine plantations, Eucalyptus sp., Ulex europaeus, Kikuyu grass. PRESERVED_SPECIMEN Starr, F. Martz, K. Collector Number: 001218-1 Maui BISH 12/18/2000
137 Hedera helix honolulu; star market garden Purchased. PRESERVED_SPECIMEN staples, g.w. collector number: 952 Oahu BISH 7/19/1994
138 Hedera helix Waimea Canyon State Park; cultivated plants from cabins near Kokee Methodist Camp. Cultivated. PRESERVED_SPECIMEN Tim Flynn 2038 Kauai PTBG 12/30/1986
139 Hedera helix East Maui, Makawao District, Piiholo, just below jct w. Olinda Rd Crawling over ground and climbing up trees to 10m ht. Associated vegetation: Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN Starr, F. Martz, K. Collector Number: 011026-1 Maui BISH 10/26/2001
140 Hedera helix W Maui, Lahaina District, Kaulalewelewe, near Haelaau Cabin Growing on and near the cabin. PRESERVED_SPECIMEN Oppenheimer, H.L. H40102 Maui BISH 4/1/2001
141 Hedera helix east maui, makawao district, piiholo, just below jct w. olinda rd Crawling over ground and climbing up trees to 10m ht. Associated vegetation: Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN starr, f. martz, k. collector number: 011026-1 Maui BISH 10/26/2001
142 Hedera helix Hawaiian Islands, Maui, E maui, Piiholo, just below jct w. Olinda Rd. Crawling on ground and climbing up trees. Associated vegetation Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN Starr, F.; Martz, K. 011026-1 K 10/26/2001
143 Hedera helix puna; volcano In Degener's tree-fern forest. PRESERVED_SPECIMEN degener, o. degener, i. collector number: 32461 Hawaii BISH 7/23/1970
144 Hedera helix w maui, lahaina district, kaulalewelewe, near haelaau cabin PRESERVED_SPECIMEN oppenheimer, h.l. collector number: h40102 Kauai BISH 4/1/2001
145 Hedera helix Hawaiian Islands, Maui, W Maui, Lahaina District, Kaulalewelewe, near Haela'au Cabin. Growing on and near the cabin. PRESERVED_SPECIMEN Oppenheimer, H. H40102 K 4/1/2001
146 Hedera helix Kaulalewelewe. On and near Haela'au Cabin and spreading into Honokowai Valley and adjacent areas. PRESERVED_SPECIMEN Hank Oppenheimer H40102 Maui PTBG 4/1/2001
147 Hedera helix Kokee. Halemanu Road near Hedychium plantings. PRESERVED_SPECIMEN Tim Flynn 968 Kauai PTBG 12/11/1984
148 Hedera helix Hawaiian Islands, Maui, W Maui, Lahaina District, Kaulalewelewe, near Haela'au Cabin. PRESERVED_SPECIMEN H. Oppenheimer H40102 US 4/1/2001
149 Hedera helix E Maui, at the top of Olinda Rd PRESERVED_SPECIMEN F. Starr & K. Martz 001218-1 Maui US 12/18/2000
150 Hedera helix hawaiian islands, maui, w maui, lahaina district, kaulalewelewe, near haela'au cabin. PRESERVED_SPECIMEN h. oppenheimer h40102 US 4/1/2001
151 Hedera helix e maui, at the top of olinda rd PRESERVED_SPECIMEN f. starr & k. martz 001218-1 Maui US 12/18/2000
152 Hedera helix e maui, piiholo, just below jct w. olinda rd PRESERVED_SPECIMEN f. starr & k. martz 011026-1 Maui US 10/26/2001
153 Hedera helix E Maui, Piiholo, just below jct w. Olinda Rd PRESERVED_SPECIMEN Starr, F.; Martz, K. 011026-1 Maui US 10/26/2001
154 Hedera helix E Maui, Piiholo, just below jct w. Olinda Rd PRESERVED_SPECIMEN F. Starr & K. Martz 011026-1 Maui US 10/26/2001
155 Hedera helix E Maui, at the top of Olinda Rd PRESERVED_SPECIMEN Starr, F.; Martz, K. 001218-1 Maui US 12/18/2000
156 Hedera helix Hawaiian Islands, Maui, W Maui, Lahaina District, Kaulalewelewe, near Haela'au Cabin. PRESERVED_SPECIMEN Oppenheimer, H. H40102 US 4/1/2001
157 Hedera helix Puna; Volcano In Degener's tree-fern forest. PRESERVED_SPECIMEN Degener, O. 32461 Hawaii BISH 7/23/1970
158 Hedera helix Captain Cook; Amy B. H. Greenwell Ethnobotanical Gardens; abandoned coffee plantation at one side of Garden In yard of house; in deep shade & also climbing shrubs & tree trunks. PRESERVED_SPECIMEN Herbst, D.R. 9303 Hawaii BISH 1/11/1991
159 Hedera helix East Maui, Makawao District, at the top of Olinda Rd Climbing in Eucalyptus trees well beyond any obvious plantings. Associated vegetation: Pine plantations, Eucalyptus sp., Ulex europaeus, Kikuyu grass. PRESERVED_SPECIMEN Starr, F. 001218-1 Maui BISH 12/18/2000
160 Hedera helix W Maui, Lahaina District, Kaulalewelewe, near Haelaau Cabin Growing on and near the cabin. PRESERVED_SPECIMEN Oppenheimer, H.L. Collector Number: H40102 Maui BISH 4/1/2001
161 Hedera helix Honolulu; Star Market Garden Purchased. PRESERVED_SPECIMEN Staples, G. 952 Oahu BISH 7/19/1994
162 Hedera helix Honolulu; Star Market Garden Purchased. PRESERVED_SPECIMEN Staples, G. 951 Oahu BISH 7/19/1994
163 Hedera helix east maui, makawao district, at the top of olinda rd Climbing in Eucalyptus trees well beyond any obvious plantings. Associated vegetation: Pine plantations, Eucalyptus sp., Ulex europaeus, Kikuyu grass. PRESERVED_SPECIMEN starr, f. martz, k. collector number: 001218-1 Maui BISH 12/18/2000
164 Hedera helix east maui, kula, old kula hwy PRESERVED_SPECIMEN starr, f. martz, k. collector number: 010419-1 Oahu BISH 4/19/2001
165 Hedera helix East Maui, Kula, old Kula Hwy PRESERVED_SPECIMEN Starr, F. Martz, K. Collector Number: 010419-1 Maui BISH 4/19/2001
166 Hedera helix captain cook; amy b. h. greenwell ethnobotanical gardens; abandoned coffee plantation at one side of garden In yard of house; sterile groundcover in deep shade & also climbing shrubs & tree trunks. PRESERVED_SPECIMEN herbst, d.r. staples iii, g.w. collector number: 9303 Hawaii BISH 1/11/1991
167 Hedera helix captain cook; amy b. h. greenwell ethnobotanical gardens Abandoned coffee plantation at one side of Garden; in yard of house. PRESERVED_SPECIMEN herbst, d.r. staples iii, g.w. collector number: 9303 Hawaii BISH 1/11/1991
168 Hedera helix east maui, kula, old kula hwy Along roadside with Syzygium cumini and Panicum maximum. PRESERVED_SPECIMEN starr, f. martz, k. collector number: 010419-1 Kauai BISH 4/19/2001
169 Hedera helix East Maui, Kula, old Kula Hwy PRESERVED_SPECIMEN Starr, F. 010419-1 Maui BISH 4/19/2001
170 Hedera helix W Maui, Lahaina District, Kaulalewelewe, near Haelaau Cabin PRESERVED_SPECIMEN H. Oppenheimer H40102 Maui HAPI 4/1/2001
171 Hedera helix 96746, Kapaa, HI, US Growing on and near the cabin. PRESERVED_SPECIMEN Oppenheimer, H. H40102 Maui HAPI 4/1/2001
172 Hedera helix East Maui, Kula, old Kula Hwy PRESERVED_SPECIMEN F. Starr & K. Martz 001218-1 Maui HAPI 12/18/2000
173 Hedera helix East Maui, Kula, old Kula Hwy Crawling on ground and climbing up trees. Associated vegetation Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN Starr, F.; Martz, K. 011026-1 Maui HAPI 10/26/2001
174 Hedera helix Oahu Island, Honolulu, University of Hawaii campus PRESERVED_SPECIMEN F. Starr & K. Martz 011026-1 Oahu HAPI 10/26/2001
175 Hedera helix East Maui, Makawao District, Piiholo, just below jct w. Olinda Rd Crawling over ground and climbing up trees to 10m ht. Associated vegetation: Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN Starr, F. 011026-1 Maui BISH 10/26/2001
176 Hedera helix honolulu; star market garden Purchased. PRESERVED_SPECIMEN staples, g.w. collector number: 951 Oahu BISH 7/19/1994
177 Hedera helix East Maui, Makawao District, at the top of Olinda Rd Climbing in Eucalyptus trees well beyond any obvious plantings. Associated vegetation: Pine plantations, Eucalyptus sp., Ulex europaeus, Kikuyu grass. PRESERVED_SPECIMEN Starr, F. Martz, K. Collector Number: 001218-1 Maui BISH 12/18/2000
178 Hedera helix honolulu; star market garden Purchased. PRESERVED_SPECIMEN staples, g.w. collector number: 952 Oahu BISH 7/19/1994
179 Hedera helix Waimea Canyon State Park; cultivated plants from cabins near Kokee Methodist Camp. Cultivated. PRESERVED_SPECIMEN Tim Flynn 2038 Kauai PTBG 12/30/1986
180 Hedera helix East Maui, Makawao District, Piiholo, just below jct w. Olinda Rd Crawling over ground and climbing up trees to 10m ht. Associated vegetation: Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN Starr, F. Martz, K. Collector Number: 011026-1 Maui BISH 10/26/2001
181 Hedera helix W Maui, Lahaina District, Kaulalewelewe, near Haelaau Cabin Growing on and near the cabin. PRESERVED_SPECIMEN Oppenheimer, H.L. H40102 Maui BISH 4/1/2001
182 Hedera helix east maui, makawao district, piiholo, just below jct w. olinda rd Crawling over ground and climbing up trees to 10m ht. Associated vegetation: Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN starr, f. martz, k. collector number: 011026-1 Maui BISH 10/26/2001
183 Hedera helix Hawaiian Islands, Maui, E maui, Piiholo, just below jct w. Olinda Rd. Crawling on ground and climbing up trees. Associated vegetation Eucalyptus spp., Pinus spp. PRESERVED_SPECIMEN Starr, F.; Martz, K. 011026-1 K 10/26/2001
184 Hedera helix puna; volcano In Degener's tree-fern forest. PRESERVED_SPECIMEN degener, o. degener, i. collector number: 32461 Hawaii BISH 7/23/1970
185 Hedera helix w maui, lahaina district, kaulalewelewe, near haelaau cabin PRESERVED_SPECIMEN oppenheimer, h.l. collector number: h40102 Kauai BISH 4/1/2001
186 Hedera helix Hawaiian Islands, Maui, W Maui, Lahaina District, Kaulalewelewe, near Haela'au Cabin. Growing on and near the cabin. PRESERVED_SPECIMEN Oppenheimer, H. H40102 K 4/1/2001
187 Hedera helix Kaulalewelewe. On and near Haela'au Cabin and spreading into Honokowai Valley and adjacent areas. PRESERVED_SPECIMEN Hank Oppenheimer H40102 Maui PTBG 4/1/2001
188 Hedera helix Kokee. Halemanu Road near Hedychium plantings. PRESERVED_SPECIMEN Tim Flynn 968 Kauai PTBG 12/11/1984
189 Hedera helix Hawaiian Islands, Maui, W Maui, Lahaina District, Kaulalewelewe, near Haela'au Cabin. PRESERVED_SPECIMEN H. Oppenheimer H40102 US 4/1/2001
190 Hedera helix E Maui, at the top of Olinda Rd PRESERVED_SPECIMEN F. Starr & K. Martz 001218-1 Maui US 12/18/2000
191 Hedera helix hawaiian islands, maui, w maui, lahaina district, kaulalewelewe, near haela'au cabin. PRESERVED_SPECIMEN h. oppenheimer h40102 US 4/1/2001
192 Hedera helix e maui, at the top of olinda rd PRESERVED_SPECIMEN f. starr & k. martz 001218-1 Maui US 12/18/2000
193 Hedera helix e maui, piiholo, just below jct w. olinda rd PRESERVED_SPECIMEN f. starr & k. martz 011026-1 Maui US 10/26/2001
194 Hedera helix E Maui, Piiholo, just below jct w. Olinda Rd PRESERVED_SPECIMEN Starr, F.; Martz, K. 011026-1 Maui US 10/26/2001
195 Hedera helix E Maui, Piiholo, just below jct w. Olinda Rd PRESERVED_SPECIMEN F. Starr & K. Martz 011026-1 Maui US 10/26/2001
196 Hedera helix E Maui, at the top of Olinda Rd PRESERVED_SPECIMEN Starr, F.; Martz, K. 001218-1 Maui US 12/18/2000
197 Hedera helix Hawaiian Islands, Maui, W Maui, Lahaina District, Kaulalewelewe, near Haela'au Cabin. PRESERVED_SPECIMEN Oppenheimer, H. H40102 US 4/1/2001
198 Hedera helix Puna; Volcano In Degener's tree-fern forest. PRESERVED_SPECIMEN Degener, O. 32461 Hawaii BISH 7/23/1970
199 Hedera helix Captain Cook; Amy B. H. Greenwell Ethnobotanical Gardens; abandoned coffee plantation at one side of Garden In yard of house; in deep shade & also climbing shrubs & tree trunks. PRESERVED_SPECIMEN Herbst, D.R. 9303 Hawaii BISH 1/11/1991
200 Hedera helix East Maui, Makawao District, at the top of Olinda Rd Climbing in Eucalyptus trees well beyond any obvious plantings. Associated vegetation: Pine plantations, Eucalyptus sp., Ulex europaeus, Kikuyu grass. PRESERVED_SPECIMEN Starr, F. 001218-1 Maui BISH 12/18/2000
201 Hedera helix W Maui, Lahaina District, Kaulalewelewe, near Haelaau Cabin Growing on and near the cabin. PRESERVED_SPECIMEN Oppenheimer, H.L. Collector Number: H40102 Maui BISH 4/1/2001
202 Hedera helix Honolulu; Star Market Garden Purchased. PRESERVED_SPECIMEN Staples, G. 952 Oahu BISH 7/19/1994
203 Hedera helix Honolulu; Star Market Garden Purchased. PRESERVED_SPECIMEN Staples, G. 951 Oahu BISH 7/19/1994
204 Hedera helix east maui, makawao district, at the top of olinda rd Climbing in Eucalyptus trees well beyond any obvious plantings. Associated vegetation: Pine plantations, Eucalyptus sp., Ulex europaeus, Kikuyu grass. PRESERVED_SPECIMEN starr, f. martz, k. collector number: 001218-1 Maui BISH 12/18/2000
205 Hedera helix east maui, kula, old kula hwy PRESERVED_SPECIMEN starr, f. martz, k. collector number: 010419-1 Oahu BISH 4/19/2001
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